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Ghost squadron is the last of two GDI military units for their Aircraft and its headquarters is located in Langley despite the Brotherhood of Nod's plan to attack both Airfields before GDI reclaimed them. Four of the A-15 Orcas were from this squadron as all others belonged to the Skull squadron. After the Liberation of Langley at the Airfields, the Orcas were used for the Battle of Alexandria where GDI attacked Nod's harbour installments to make way for a special landing by GDI Hovercrafts carrying two Mammoth tanks on the north-western side of the map. They finally destroyed Nod's three compounds and the base before heading for Egypt's captial city, Cairo to destroy the Cairo Nuclear Launch Facility that was responsible for starting the Third Tiberium War.

Skull squadron is one of two GDI military units and taking charge of its aircraft. Four of the A-15 Orcas were from this squadron as the other four belonged to the Ghost squadron. Their headquarters is located at an Airfield where Nod forces stood their hold and started attacking the two Airfields at Langley Air Force Base. When GDI reclaimed the Airfields, the squadron reunited at this base and attacked Nod's primary garrison by destroying key structures prior to the final objective. Orcas were now deployed at Alexandria in Egypt where they suddenly destroyed two Battleships and loading cranes which are in the hands of Nod with three compounds for controlling the city's region in a dangerous Red Zone.

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